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BiologyCollege of Arts and Sciences Chairperson Departmental E-mail Departmental URL Graduate Faculty Graduate Faculty(An asterisk [*] denotes associate membership in University Graduate School faculty.) Clyde Culbertson Professor Distinguished Professors Professors Associate Professors Assistant Professors Senior Scientist Assistant Scientist Adjunct Professors Adjunct Associate Professors Adjunct Assistant Professors Director of Graduate Studies Degrees OfferedMaster of Arts and Doctor of Philosophy in ecology and evolutionary biology; Doctor of Philosophy in genetics; Master of Arts and Doctor of Philosophy in microbiology; Doctor of Philosophy in molecular, cellular, and developmental biology; Master of Arts and Doctor of Philosophy in plant sciences; Master of Arts and Doctor of Philosophy in zoology; and Master of Arts for Teachers. Special Departmental Requirements(See also general University Graduate School requirements.) Admission Requirements Special Requirement for the M.A. Degree Ph.D. Qualifying Examination Final Examination Other Provisions Ecology and Evolutionary BiologyMaster of Arts DegreeCourse Requirements Thesis Final Examination Doctor of Philosophy Degree Course Requirements Concentration Area RequirementsEcology/Population BiologyE455 (SPEA) Limnology L575 Biodiversity and Ecosystem Functioning L578 Advanced Population Biology L579 Community Ecology L591 Plant Population Biology—An Experimental Approach Evolutionary Biology Behavior/Physiology Minor Foreign Language/Research Skill Requirements Final Examination GeneticsMolecular, Cellular, and Developmental BiologyDoctor of Philosophy DegreePrograms leading to the Ph.D. degrees in genetics, and in molecular, cellular, and developmental biology are administered by the Faculty Committee on Molecular Biology and Genetics (MBG), in collaboration with members of the Department of Chemistry. The Ph.D. in plant sciences can be pursued under the supervision of MBG or that of ecology and evolutionary biology, depending upon the nature of a student's research interests. Programs leading to the Ph.D. degrees in genetics, and in molecular, cellular, and developmental biology are administered by the Faculty Committee on Molecular Biology and Genetics (MBG), in collaboration with members of the Department of Chemistry. The Ph.D. in plant sciences can be pursued under the supervision of MBG or that of ecology and evolutionary biology, depending upon the nature of a student's research interests. Common Requirements At the end of the first year, each student selects a research advisor and laboratory. Together with the advisor, the student also selects the other members of an advisory committee of three or four faculty members appropriate to the student's intended degree and one from the prospective minor field (see below). This advisory committee guides and monitors the student's subsequent independent work and guides the student's selection of advanced courses. MBG requires that each student meet with the advisory committee at least once per year. The MBG-administered degree programs require a total of 90 credit hours including the core program and at least three advanced courses (see below). They also require that each student register for Journal Club (Z620) during the second year and make two oral presentations in Journal Club during the course of his/her career. Each student must also take a Z620 Grant Writing course in their second year, and a Z620 course in Scientific Practices (Ethics) during the third year, and each student must teach for at least two semesters. Grades Preliminary Examination Satisfactory Progress Toward a Degree Thesis Advanced Courses and Minor Each student must select a minor field distinct from the chosen degree. Ordinarily a student will select as a minor one of the MBG degree programs not selected for the major. In those cases, the Core Program courses meet minor requirements. In some cases a student may select another minor and must meet any additional requirements set by that minor. For students from other programs who wish to minor in one of the MBG degree areas, the requirement is 6 credit hours of work in that field. The course selection should be approved by the director of MBG. MicrobiologyDegree programs are available for students with interests in many areas of microbiology. Each student's curriculum is designed by the student in consultation with the graduate program director, the student's mentor, and an appointed advisory committee. Master of Arts Degree with a Research ThesisCourse Requirements Grades Thesis Final Examination Master of Arts Degree with a Library ThesisThe department also offers a program in microbiology leading to a terminal master's degree that does not require a laboratory research project. A student enrolled in this program will write a thesis critically evaluating and reviewing some aspect of microbiology reported in the literature. All other requirements for the degree are identical to those stated above for the research-thesis Master of Arts. The degree is designed to give individuals an opportunity to pursue graduate study at the master's level without acquiring expertise in laboratory research. Doctor of Philosophy DegreeCourse Requirements Grades Advisory Committee Thesis Final Examination Plant SciencesMaster of Arts DegreeCourse Requirements Grades Thesis Doctor of Philosophy DegreePh.D. students choosing a molecular approach will follow the procedures in all respects (courses, seminars, research rotations, preliminary examination, etc., of the Genetics and MCDB (Molecular, Cellular, and Developmental Biology) graduate programs. Likewise, students choosing an organismal approach will follow the exact procedures of the Ecology and Evolutionary Biology program. ZoologyEach degree program is tailored to the specific interests and needs of the student. Master of Arts DegreeCourse Requirements Grades Thesis Doctor of Philosophy DegreeCourse Requirements Minor Master of Arts for Teachers DegreeThe Master of Arts for Teachers in biology is offered by the University Graduate School (not the School of Education) to provide training beyond the bachelor's degree for those who intend to teach in junior or senior high school and who wish additional training in biology. Each student in the program must possess a teacher's certificate by the time the degree is conferred, with the exception of international students who intend to return to their native country. Admission Requirement Course Requirements Certification Requirements CoursesM300 Biomedical Sciences Documentation (1 cr.) M315 Microbiology Laboratory (2 cr.) M500 Introduction to Research (Microbiology) (1-6 cr.) P: graduate standing. Objectives and techniques of microbiological research. Assignment to a research problem with a faculty member to be completed in two semesters. T500 Project Laboratory in Biotechnology (6 cr.) In this course students explore the different stages of scientific investigation by performing research using the techniques of chemistry, biochemistry, molecular biology, genetics, and cell biology on problems related to biotechnology. Students design and execute research projects under supervision of the instructor in a teaching laboratory setting on problems chosen in consultation with the instructor. L501 Independent Study (1-6 cr.) P: written consent of faculty member supervising work. Supervised work. S/F grading. T501 Topics in Biotechnology I (2 cr.) Students read and analyze research articles from the current literature and present the articles in a journal club format. Students will practice their presentation with the instructor prior to presenting to the group and will receive feedback on the content and the presentation style. Guest lecturers from industry are invited to present on a wide range of topics relevant to biotechnology. T502 Topics in Biotechnology II (2 cr.) Follows from BIOL T501. Students read and analyze research articles from the current literature and present the articles in a journal club format. Articles can cover any area of biotechnology or any area relevant to biotechnology. Occasionally, invited guest lecturers from industry are invited to present on a wide range of topics relevant to biotechnology. L505 Evolution of Development (3 cr.) P: senior or graduate standing and consent of instructor. An integrative approach to the link between development and the evolution of morphology. Topics: evolution of developmental mechanisms and of developmental regulatory genes, production of evolutionary changes through changes in developmental processes, developmental constraints, and origins of major body plans. Z508 Advanced Ornithology (4 cr.) P: Z406. Emphasis on avian ecology, distribution, and behavior; discussion and evaluation of recent literature. Field work includes investigation of populations of a wintering species and a breeding species. L509 Field Exercises for Biology Education (1-5 cr.) L509 is a graduate course for students in biology and education with an intended career in biology education. Credits are variable (1-5) and will be arranged. Students will design field exercises based at the Indiana University Research and Teaching Preserve on topics in organismal biology and ecology appropriate for public school and other outside groups. L510 Introduction to the Research Laboratory (3 cr.) P: graduate standing. Objectives and techniques of biological research. Completion of a one-semester research problem with a faculty member. T510 Theory and Applications of Biotechnology Lecture I (3 cr.) This advanced, graduate-level course will focus on the applications of molecular genetics and recombinant DNA in biotechnology. Fundamental concepts of relevant molecular biology and biochemistry will be covered in depth in the first portion of the class, followed by sections on recombinant DNA technology, macromolecular purification and genomics/bioinformatics. M511 Molecular Biology of Prokaryotes (3 cr.) P: CHEM C584. The course will first develop an understanding of nucleic acid structure and function to a professional level, then use these principles to explore molecular aspects of gene expression and evolution. Emphasis will be on prokaryotes. T511 Theory and Application of Biotechnology Lecture II (3 cr.) Course continues from BIOL T510, Theory and Applications of Biotechnology Lecture I, but focuses on applications of biotechnology including genetic engineering of plants and animals, bioremediation, biopharmaceutical production, vaccine development, and molecular diagnostics. Bioengineering principles of fermentation, scale-up and high throughput functional screening will be an important component of this material. M512 Molecular Biology of AIDS Virus (3 cr.) P: CHEM C341 and BIOL L311. A detailed consideration of the human immunodeficiency virus (HIV, causative agent of AIDS). The functions of the HIV genes and how those functions affect pathology and normal cellular mechanisms. T515 Theory and Applications of Biotechnology Laboratory I (3 cr.) Students will learn advanced laboratory techniques that are currently used in biotechnology. This course is designed to cover advanced techniques at a deep level. As far as possible the laboratory exercises will be coordinated with BIOL T510, Theory and Applications of Biotechnology Lecture I. There will be two modules, one emphasizing cell biology and one emphasizing molecular biology. T516 Theory and Applications of Biotechnology Laboratory II (3 cr.) Course continues from BIOL T515 Theory and Applications of Biotechnology Laboratory I. As far as possible the laboratory exercises will be coordinated with BIOL T511, Theory and Applications of Biotechnology Lecture II. There will be two modules, one emphasizing cell biology and one emphasizing molecular biology. L519 Bioinformatics: Theory and Application (3 cr.) Overview of theory and applications in bioinformatics, based on fundamentals of molecular biology and information sciences. Common problems, data and tools in the field are outlined. These include biosequence analysis, alignment and assembly, genomics, proteomics and phylogenetics, biological databases and data mining, and internet bio-information services. L520 Seminar in Genetics (cr. arr.) P: L364 or Z420 or equivalents. L521 Problems in Genetics-Higher Organisms (3 cr.) P: L364 or equivalent. Selected topics in the genetics of higher organisms emphasizing studies at the molecular level. T521 Research Design and Ethics (2 cr.) Fundamentals of research protocol design and planning with applications to practical problems. Problems of research ethics and the role of biotechnology in human society will be addressed in class discussion and seminars. L522 Advanced Eukaryotic Molecular Genetics (3 cr.) P: consent of instructor; beginning course in genetics. Correlation of genetic data with changes in chromosome structure and number. Mechanics of chromosome behavior in crossing over and disjunction. L523 Critical Analysis of the Scientific Literature (1-6 cr.) Detailed analysis of current research papers in biology. Emphasis on experimental design, research methods, interpretation of results, and suitability of controls. Generally taken in the first semester of graduate residence. Topics may vary to suit specific fields (e.g., molecular, cellular, and developmental biology and genetics, or ecological and evolutionary biology). M525 Topics in Microbial Biochemistry and Physiology (3 cr.) P: graduate standing and C483 or M350 or equivalent. The course will consider topics in physiology and biochemistry of eukaryotic and prokaryotic microorganisms. Subjects include membrane physiology and regulatory networks in metabolism and gene expression. L529 Bioinformatics in Molecular Biology and Genetics: Practical Applications (4 cr.) P: I501, I502, L519, or consent of instructor. Practical experience in a range of data analysis and software engineering methods applied to molecular biology data. B530 Anatomy and Morphology Seminar (cr. arr.) P: consent of instructor. Seminars will include current research studies in plant anatomy and morphology. M540 Medical Microbiology and Medical Immunology (2-5 cr.) Basic concepts of immunology; microorganisms as agents of disease; host-parasite relationships; epidemiology; chemotherapy. Z540 Genetics of Populations (3 cr.) P: consent of instructor: R: Z465, MATH M216 or equivalent. Survey of the theoretical basis of population genetics and a review of current problems and experimental findings. Content varies from year to year. M545 Medical Microbiology Laboratory (1 cr.) P: M540. Laboratory experiments to illustrate material discussed in M540. M550 Microbiology (3 cr.) P: two semesters of college chemistry; L211 recommended prior or concurrently. Application of fundamental principles to the study of microorganisms. Significance of microorganisms to humans and their environment. Critical evaluation of current microbiological literature. B555 Special Topics in Plant Systematics (3 cr.) Topics vary from year to year. Examples of subjects to be treated: phylogeny and families of flowering plants, biology of ferns, biosystematics, molecular markers in populational biology and systematics. Enrollment of advanced undergraduates encouraged. L555 Alternative Approaches to Teaching College Biology (2 cr.) Frameworks for teaching college biology. Addresses different teaching objectives (knowledge, applications, scientific thinking, ethical and policy considerations); different teaching methods (lectures, readings, recitations, discussions, exercises, experiments, projects); student heterogeneity (expectations, abilities, development, learning styles); evaluation and grading; course and curriculum design; and evaluation and improvement of teaching. B560 Seminar in Systematics (cr. arr.) P: consent of instructor. Topics vary each semester. L560 Physiological Ecology (3 cr.) Influence of the abiotic environment on energy and material transfers in individual organisms, with emphasis on terrestrial animals. Z566 Laboratory in Endocrinology (2 cr.) P: Z466. Development and structure of major endocrine glands; their role in maintaining constancy of internal environment. Limited to 12 students. L567 Evolution (3 cr.) P: graduate standing in Psychology or Biology or consent of the instructor. Topics include quantitative genetics, population genetics and strategic models of natural selection. Special topics include: life history theory, sex and sexual selection, kin selection, shifting-balance theory, speciation, macroevolution, and comparative methods. B570 Seminar in Physiology and Molecular Biology of Plants (cr. arr.) P: consent of instructor. L570 Seminar in Ecology and Environmental Biology (1 cr.) P: consent of instructor. Presentations and discussions of current research in evolution, ecology, and behavior. May be repeated for credit. B572 Photobiology (3 cr.) P: S305 or L367 or CHEM C483 or equivalent. Biochemical and biophysical relationship between light and biological systems. Topics will include photosynthesis, visual processes, photorespiration, phototaxis, bioluminescence, and photomorphogenesis, with emphasis on photosynthesis. L572 Microbial Ecology (3 cr.) Principles of microbial ecology with emphasis on the population, community, and ecosystem ecology of bacteria and fungi. B573 Special Topics in Plant Physiology (2-5 cr.) P: consent of instructor. Advanced topics in plant physiology. With consent of instructor, may be taken more than once for credit. L575 Ecosystem Structure and Function (3 cr.) P: L473 and L474 (or equivalent) or instructor's consent. Does biodiversity matter? Analysis of relationships between biodiversity and ecosystem functioning. Emphasis on current literature, including theoretical and empirical work. Lectures will alternate with class discussion and debate. M575 Human Parasitology (4 cr.) P: BIOL M310 and M315. Biology of human parasites focusing on their etiology, epidemiology, immunology, diagnosis, and treatment. Major groups of protozoa, helminths, and medically important arthropods covered. Independent research assigned on a special topic. Lab presents both live and fixed materials complementing lecture. B576 Developmental Plant Physiology (3 cr.) P: consent of instructor. Chemically oriented; examination of substances uniquely involved in growth and development in higher plants. Application of information to lower plants only briefly discussed. Z576 Invertebrate Zoology Laboratory (2 cr.) P or C: Z374. Laboratory and field studies of invertebrates, with an emphasis on experiments with living specimens. B577 Plant Biochemistry (2 cr.) A comparative treatment of selected biochemical topics, emphasizing unique or important processes in plant metabolism and development. L578 Advanced Population Biology (3 cr.) P: courses in ecology, genetics, and basic calculus, and permission of instructor. A detailed assessment of population-ecological and population-genetic theory, and the factors determining the size and composition of animal populations in nature. L579 Community Ecology (3 cr.) P: ecology and genetics. Survey of ecological and evolutionary topics between population and ecosystem levels. Review of scientific levels of selection and speciation. Major emphasis on interactions among populations (consumer-producer, competition, symbiosis, etc.) and community analysis (island biogeography, niche, diversity, and community structure). L580 Introduction to Research (1 cr.) Individual faculty from the various graduate programs in biology present seminars on their research programs. Discussion between students and faculty about possible thesis research projects is encouraged. L581 Behavioral Ecology (3 cr.) Integrated elements of ethology, physiology, ecology, and evolutionary biology providing a synthetic approach to animal behavior. Emphasis on integrated studies providing new insights into both evolutionary and mechanistic questions. Students are asked to analyze the literature critically and debate controversial issues actively. L585 Molecular Genetics (3 cr.) The molecular basis of genetic interactions, with emphasis on microbial systems. The course covers the molecular mechanisms of mutation, suppression, recombination, complementation, etc., as well as mechanisms for gene transfer in bacteria and bacteriophage. The application of genetic analysis to variety of molecular biological topics is emphasized. L586 Cell Biology (4.5 cr.) Critical analysis of recent advances in our understanding of molecular organization and function of cellular structures. The emphasis of this course will be on eukaryotic cells. Topics include membrane organization, cytoskeleton assembly and functions, signal transduction, cell-cycle regulation, protein sorting, and vesicle trafficking. L587 Developmental Biology (4.5 cr.) Evaluation of classical and current molecular and genetic approaches to studying development of eukaryotic organisms. A significant portion of the course is devoted to discussing recent findings from molecular genetic studies in Drosophila and C. elegans. L590 Seminar in Molecular, Cellular, and Developmental Biology (2 cr.) P: consent of instructor. Presentation and discussion of topics in molecular and cellular biology as seminar by students. Topics from current literature. Concentration on a particular area each semester to be announced before registration. S/F grading. L591 Plant Population Biology—An Experimental Approach (3 cr.) P: ecology course and evolution course. The mechanisms by which plants, as individuals, contribute to development of population structure. Experimental studies of intra- and inter-specific mechanisms of population regulation, reproduction, and vegetative growth. Emphasis on development and physiological characteristics which determine mode of interaction. Greenhouse projects designed and conducted by students. L600 Special Topics in Genetics (cr. arr.) P: L364 or equivalent. Topics not extensively treated in other courses, e.g., population genetics, human genetics, immunogenetics, biochemical genetics of clones of mammalian cells. Topic presented will not be duplicated within three to five years. L600 carries credit in Plant Sciences, Microbiology, and Zoology programs. M610 Recent Advances in Microbiology (1-3 cr.) P: graduate standing in microbiology or related area. Course content changes each semester so that over a cycle of several years, major research areas are covered. May be repeated for credit. M612 Microbial Development (3 cr.) P: graduate standing or consent of instructor. An analysis of recent publications concerned with the biochemistry of development in viral, prokaryotic, and simple eukaryotic systems. The topics vary and emphasize the regulatory aspects of development. Cell differentiation and cell-cell interactions are discussed. Z620 Special Topics in Zoology (cr. arr.) P: advanced undergraduate or graduate standing. Topics not extensively treated in other courses, e.g., theoretical zoology, oceanography, reservoir limnology, human ecology, biochemistry, viruses and disease, critical analysis of the scientific literature, and other fields. Topics presented will be treated every three to five years. L800 Research (1-15 cr.)
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